/* Figma tsWYlPm3nOMps1SkY9NOFq / Site redesign 2136:3561, read 2026-10-31.
   Shared by the server reading pages and the public React application. */
.site-public-app, .site-public-navigation, body:has(> .reader-header) {
  --site-canvas: var(--eco-primitive-neutral-black, #101010);
  --site-panel: var(--eco-primitive-neutral-charcoal, #222222);
  --site-feature: var(--eco-primitive-brand-olive-graphite, #2e302c);
  --site-accent: var(--eco-primitive-brand-moss, #4f603c);
  --site-text: var(--eco-primitive-neutral-white, #fff);
  --site-muted: var(--eco-primitive-neutral-light-gray, #acacac);
  --site-line: var(--eco-primitive-neutral-graphite, #434343);
  --eco-semantic-background-base: var(--site-canvas);
  --eco-semantic-background-navigation: var(--site-panel);
  --eco-semantic-background-header: var(--site-panel);
  --eco-semantic-surface-primary: var(--site-panel);
  --eco-component-product-sidebar-background: var(--site-panel);
  --eco-component-product-header-background: var(--site-panel);
  --bg: var(--site-canvas); --panel: var(--site-panel); --panel2: var(--site-feature);
  --ink: var(--site-text); --muted: var(--site-muted); --line: var(--site-line);
  background: var(--site-canvas); color: var(--site-text);
  font-family: var(--eco-font-family-sans, Geist, sans-serif);
}
:root[data-eg-theme="light"] .site-public-app, :root[data-eg-theme="light"] .site-public-navigation, :root[data-eg-theme="light"] body:has(> .reader-header) {
  --site-canvas: var(--eco-primitive-neutral-white); --site-panel: #f5f5f3;
  --site-feature: #e9ede4; --site-text: var(--eco-primitive-neutral-black);
  --site-muted: var(--eco-primitive-neutral-gray); --site-line: #d7d9d3;
}
.site-design-header, .reader-header.site-design-header {
  position: fixed; inset: 16px 16px auto; z-index: 1200; box-sizing: border-box;
  height: 56px; min-height: 56px; margin: 0; padding: 8px 12px; gap: 16px;
  display: flex; align-items: center; flex-wrap: nowrap; border: 0; border-radius: 8px;
  background: var(--site-panel); color: var(--site-text); transition: transform .18s ease;
}
[data-header-hidden="true"] .site-design-header { transform: translateY(-88px); }
.site-header-brand { display: flex; align-items: center; gap: 24px; flex: 0 0 244px; }
.site-wordmark, .reader-header .brand .site-wordmark { display: block; width: 176px; height: 23.079px; }
.site-icon-button, .site-design-header .reader-desktop-menu, .site-design-header .reader-menu > summary {
  display: inline-flex; align-items: center; justify-content: center; width: 40px; height: 40px;
  min-height: 40px; padding: 0; border: 0; border-radius: 8px;
  color: var(--site-text); background: var(--eco-primitive-neutral-graphite); flex: 0 0 40px;
}
.site-global-search, .site-design-header .reader-header-search {
  display: flex; align-items: center; flex: 0 1 624px; width: 624px; max-width: 624px; min-width: 120px;
  height: 40px; margin: 0 0 0 auto; padding: 0 12px; gap: 4px; border: 0;
  border-radius: 8px; background: var(--eco-primitive-neutral-gray);
}
.site-global-search input, .site-design-header .reader-header-search input {
  flex: 1; width: 0; min-width: 0; height: 40px; padding: 0; margin: 0; border: 0; outline-offset: 2px;
  background: transparent; color: var(--eco-primitive-neutral-white); font: inherit; font-size: 12px;
}
.site-global-search input::placeholder, .site-design-header .reader-header-search input::placeholder { color: var(--eco-primitive-neutral-white); opacity: .9; }
.site-global-search button, .site-design-header .reader-header-search button {
  display: flex; align-items: center; justify-content: center; flex: 0 0 20px;
  width: 20px; min-height: 20px; height: 20px; padding: 0; border: 0; color: white; background: transparent;
}
.site-header-help { position: relative; background: transparent; text-decoration: none; }
.site-header-help span { position: absolute; font-size: 13px; line-height: 18px; color: var(--site-muted); }
.site-languages.reader-languages { display: flex; flex: 0 0 88px; gap: 0; height: 40px; padding: 4px; box-sizing: border-box; border: 0; border-radius: 9px; background: var(--eco-primitive-neutral-graphite); }
.site-languages a { display: flex; justify-content: center; align-items: center; width: 40px; min-height: 32px; border-radius: 7px; color: #acacac; font-size: 13px; text-decoration: none; }
.site-languages a[aria-current="true"] { color: white; background: var(--eco-primitive-neutral-gray); }
.site-header-account, .site-design-header .reader-header-login, .site-design-header [data-reader-account="header"] a {
  display: inline-flex; align-items: center; justify-content: center; box-sizing: border-box;
  width: 112px; min-height: 40px; padding: 0 12px; border: 0; border-radius: 8px;
  background: var(--eco-primitive-neutral-graphite); color: #fff; font-size: 14px; font-weight: 500; text-decoration: none; white-space: nowrap;
}
.site-public-app [data-testid="app-sidebar"], .platform-sidebar {
  background: var(--site-panel); border: 0; border-radius: 16px;
}
.site-public-navigation > nav { padding: 16px; scrollbar-gutter: auto; }
.site-navigation { display: flex; flex-direction: column; gap: 12px; }
.site-nav-group, .platform-sidebar .site-nav-group { margin: 0; padding: 0; border: 0; }
.site-nav-group h2, .platform-sidebar .site-nav-group h2 { margin: 0 0 4px; padding: 4px 8px; color: var(--site-muted); font-size: 12px; line-height: 16px; font-weight: 400; text-transform: none; letter-spacing: 0; }
.site-nav-link, .platform-sidebar .site-nav-link {
  display: flex; align-items: center; gap: 8px; box-sizing: border-box; min-width: 0;
  height: 28px; min-height: 28px; padding: 0 8px; margin: 0; border: 0; border-radius: 8px;
  color: var(--site-muted); text-decoration: none; font-size: 14px; line-height: 16px; font-weight: 400;
}
.site-nav-link strong { font-weight: 400; }
.site-nav-link[aria-current="page"], .platform-sidebar .site-nav-link[aria-current="page"] { background: var(--eco-primitive-neutral-gray); color: white; }
.site-nav-link:hover { background: var(--site-line); color: var(--site-text); }
.site-nav-home { height: 32px; padding-left: 16px; }
.site-nav-icon { display: inline-flex; align-items: center; justify-content: center; width: 20px; height: 20px; flex: 0 0 20px; }
.site-nav-icon img { display: block; max-width: 20px; max-height: 20px; }
.site-navigation--compact .site-nav-link { height: auto; min-height: 56px; flex-direction: column; padding: 6px 0; gap: 4px; text-align: center; font-size: 10px; }
.site-design-footer { display: flex; justify-content: space-between; align-items: center; gap: 24px; margin-top: 48px; padding: 24px 0 32px; border-top: 1px solid var(--site-line); color: var(--site-muted); font-size: 14px; line-height: 20px; }
.site-design-footer nav { display: flex; align-items: center; gap: 32px; flex-wrap: wrap; }
.site-design-footer a { display: inline-flex; min-height: 44px; align-items: center; color: inherit; text-decoration: none; }
.site-design-footer a:hover { color: var(--site-text); text-decoration: underline; }
.site-section { margin: 0; padding: 32px 0; border-top: 1px solid var(--site-line); }
.site-eyebrow { font-size: 10px; line-height: 1.35; color: var(--site-muted); text-transform: uppercase; letter-spacing: .02em; }
.site-reading-tabs { display: flex; gap: 8px; margin: 24px 0; flex-wrap: wrap; border-bottom: 1px solid var(--site-line); padding-bottom: 16px; }
.site-reading-tabs a { display: inline-flex; align-items: center; min-height: 40px; padding: 0 16px; color: var(--site-text); background: var(--site-panel); border-radius: 8px; text-decoration: none; font-size: 13px; }
.site-reading-tabs a:hover { background: var(--site-accent); }
.site-article-page .scientific-document { background: none; border: 0; padding: 0; }
.site-article-page .scientific-header { margin: 0 0 32px; padding: 0 0 24px; border-bottom: 1px solid var(--site-line); }
.site-article-page .scientific-header h1, .site-article-page .scientific-header h2 { margin: 0; font-size: 38px; line-height: 1.12; font-weight: 500; }
.site-article-page .scientific-layout { display: grid; grid-template-columns: minmax(0, 1fr) 240px; gap: 32px; }
.site-article-page .scientific-toc-desktop { grid-column: 2; grid-row: 1; align-self: start; background: var(--site-panel); padding: 24px; border-radius: 12px; }
.site-article-page .scientific-body { grid-column: 1; grid-row: 1; min-width: 0; }
.site-article-page .scientific-lead { background: var(--site-feature); padding: 24px; border-radius: 12px; }
.site-article-page .scientific-body > section { scroll-margin-top: 104px; }
.article-connections > p { color: var(--site-muted); font-size: 14px; line-height: 1.5; }
.connected-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 16px; margin-top: 24px; }
.connected-card { display: flex; flex-direction: column; min-width: 0; padding: 24px; border: 1px solid transparent; border-radius: 12px; background: var(--site-panel); color: var(--site-text); text-decoration: none; }
.connected-card:nth-child(3n + 2) { background: var(--site-feature); }
.connected-card h3 { margin: 16px 0 8px; font-size: 18px; line-height: 1.35; font-weight: 500; }
.connected-card p { margin: 0 0 32px; color: var(--site-muted); font-size: 14px; line-height: 1.5; }
.connected-card-action { margin-top: auto; padding-top: 12px; border-top: 1px solid var(--site-line); color: var(--eco-semantic-text-brand); font-size: 12px; }
.connected-card:hover { border-color: var(--site-accent); }
.site-article-page .scientific-body .connected-grid { grid-template-columns: repeat(2, minmax(0, 1fr)); }
.site-public-app :focus-visible, body:has(>.reader-header) :focus-visible { outline: 2px solid var(--eco-semantic-focus-ring, #acacac); outline-offset: 4px; }
@media (min-width: 1024px) {
  .reader-header.site-design-header + .platform-sidebar { top: 88px; left: 16px; width: 272px; padding: 16px; box-sizing: border-box; max-height: calc(100dvh - 104px); overflow-y: auto; }
  .platform-sidebar > nav { display: flex; flex-direction: column; gap: 12px; }
  body:has(>.site-design-header) > main { margin: 100px 16px 0 304px; width: auto; max-width: none; padding: 0; }
  body:has(>.site-design-header) > .site-design-footer { margin-left: 304px; margin-right: 16px; max-width: none; }
  :root[data-navigation-expanded="false"] body:has(>.site-design-header) > main,
  :root[data-navigation-expanded="false"] body:has(>.site-design-header) > .site-design-footer { margin-left: 116px; }
  :root[data-navigation-expanded="false"] .reader-header.site-design-header + .platform-sidebar { width: 84px; padding: 8px; }
  :root[data-navigation-hidden="true"] body:has(>.site-design-header) > main,
  :root[data-navigation-hidden="true"] body:has(>.site-design-header) > .site-design-footer { margin-left: 16px; }
  .site-design-header .reader-menu { display: none; }
}
@media (max-width: 1150px) { .site-header-brand { flex-basis: auto; gap: 16px; } .site-design-header { gap: 8px; } .site-header-help { display: none; } }
@media (max-width: 1023px) {
  .site-design-header, .reader-header.site-design-header { inset: 8px 8px auto; padding: 8px; }
  .site-header-brand { gap: 8px; } .site-wordmark, .reader-header .brand .site-wordmark { width: 132px; height: 17.31px; }
  .site-design-header .reader-desktop-menu { display: none; }
  body:has(>.site-design-header) > main { margin: 88px 16px 0; padding: 0; width: auto; }
  body:has(>.site-design-header) > .site-design-footer { margin: 32px 16px 0; }
  .site-article-page .scientific-layout { grid-template-columns: minmax(0, 1fr); }
  .site-article-page .scientific-toc-desktop { display: none; }
  .connected-grid { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .site-design-footer { align-items: flex-start; flex-direction: column; }
}
@media (max-width: 650px) {
  .site-design-header, .reader-header.site-design-header { gap: 8px; } .site-global-search, .site-design-header .reader-header-search { flex: 1 1 40px; width: 40px; min-width: 40px; padding: 0 10px; }
  .site-global-search input, .site-design-header .reader-header-search input { position: absolute; width: 1px; height: 1px; overflow: hidden; clip-path: inset(50%); }
  .site-global-search:focus-within, .site-design-header .reader-header-search:focus-within { position: absolute; left: 8px; right: 8px; width: auto; z-index: 2; }
  .site-global-search:focus-within input, .site-design-header .reader-header-search:focus-within input { position: static; width: 0; height: 40px; clip-path: none; }
  .site-header-account, .site-design-header .reader-header-login, .site-design-header [data-reader-account="header"] a { width: auto; min-width: 56px; font-size: 12px; padding: 0 8px; }
  .site-languages.reader-languages { flex-basis: 64px; width: 64px; } .site-languages a { width: 28px; font-size: 12px; }
  .site-article-page .scientific-header h1, .site-article-page .scientific-header h2 { font-size: 30px; }
  .connected-grid, .site-article-page .scientific-body .connected-grid { grid-template-columns: minmax(0, 1fr); }
  .connected-card { padding: 20px; } .site-reading-tabs a { padding: 0 12px; }
  .site-design-footer nav { gap: 8px 24px; }
}
@media (prefers-reduced-motion: reduce) { .site-design-header { transition: none; } }

/* Home, using the same composition for server HTML and React. */
.site-home h1, .site-home h2, .site-home h3 { color: var(--site-text); font-weight: 500; letter-spacing: -.015em; }
.site-home-heading { margin: 0 0 48px; }
.site-home-heading h1 { margin: 0 0 12px; font-size: 36px; line-height: 1.15; }
.site-home-heading p { margin: 0; color: var(--site-muted); font-size: 16px; line-height: 1.5; }
.site-home h2 { margin: 0 0 24px; font-size: 20px; line-height: 1.3; }
.site-home h3 { margin: 12px 0; font-size: 20px; line-height: 1.25; }
.site-home .site-section { padding: 36px 0 48px; }
.site-home-entry { margin-bottom: 48px; }
.site-home-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 20px; }
.site-home-card { display: flex; flex-direction: column; min-width: 0; min-height: 192px; padding: 20px; box-sizing: border-box; border-radius: 12px; background: var(--site-feature); text-decoration: none; color: var(--site-text); }
.site-home-card p { margin: 0 0 24px; color: var(--site-muted); font-size: 14px; line-height: 1.45; }
.site-home-card .site-eyebrow { color: var(--eco-semantic-text-brand); }
.site-home-card .connected-card-action { font-size: 12px; }
.site-home-card:hover, .site-home-anatomy:hover { box-shadow: inset 0 0 0 1px var(--site-accent); }
.site-home-groups { grid-template-columns: repeat(4, minmax(0, 1fr)); gap: 24px; }
.site-home-groups .site-home-card { background: var(--site-panel); min-height: 172px; }
.site-home-groups h3 { font-size: 18px; }
.site-home-feature { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 36px; align-items: center; }
.site-home-feature figure { margin: 0; min-width: 0; }
.site-home-feature figure img { display: block; width: 100%; aspect-ratio: 540 / 330; height: auto; object-fit: cover; border-radius: 12px; }
.site-home-feature figcaption { margin-top: 8px; font-size: 11px; line-height: 1.4; color: var(--site-muted); }
.site-home-feature h3 { margin: 0 0 16px; font-size: 30px; }
.site-home-feature p { margin: 0; max-width: 480px; font-size: 16px; line-height: 1.5; color: var(--site-muted); }
.site-home-actions { display: flex; flex-wrap: wrap; gap: 16px; margin-top: 48px; }
.site-home .primary, .site-home .secondary { display: inline-flex; align-items: center; justify-content: center; min-height: 44px; box-sizing: border-box; min-width: 180px; padding: 12px 24px; border-radius: 8px; font-size: 14px; font-weight: 500; text-decoration: none; color: var(--site-text); }
.site-home .primary { background: var(--site-accent); border: 1px solid var(--site-accent); }
.site-home .secondary { background: var(--site-panel); border: 1px solid var(--site-accent); }
.site-home-research { display: grid; grid-template-columns: 1.2fr 1fr; gap: 32px; }
.site-home-research > .site-home-card { background: var(--site-panel); }
.site-home-anatomy { position: relative; display: flex; flex-direction: column; min-height: 292px; padding: 20px 24px; border-radius: 12px; box-sizing: border-box; overflow: hidden; background: #384a2d; color: #a9c58c; text-decoration: none; }
.site-home-anatomy > .site-eyebrow { color: inherit; }
.site-home-anatomy > span:nth-last-child(2) { margin-top: auto; padding-top: 210px; font-size: 12px; text-transform: uppercase; z-index: 1; }
.site-home-anatomy > small { font-size: 11px; color: #a7aba3; z-index: 1; }
.site-anatomy-rings { position: absolute; inset: 0; display: flex; align-items: center; justify-content: center; pointer-events: none; }
.site-anatomy-rings img { position: absolute; max-width: 90%; }
.site-home-account { min-height: 172px; }

/* Existing attributed passport data keeps its reading order and anchors. */
.species-passport { border: 0; background: transparent; }
.passport-title { margin: 32px 0 24px; align-items: flex-start; }
.passport-title h1 { font-size: 38px; line-height: 1.15; font-weight: 500; }
.passport-tabs { border-bottom: 1px solid var(--site-line); gap: 24px; }
.passport-tabs a { min-height: 44px; font-size: 14px; }
.passport-intro { grid-template-columns: minmax(0, 1fr) minmax(0, 1.05fr); gap: 24px; margin: 32px 0; }
.passport-photo, .passport-overview { border-radius: 12px; overflow: hidden; }
.passport-overview { padding: 24px; background: var(--site-feature); border: 0; }
.passport-overview h2 { font-size: 28px; font-weight: 500; }
.passport-overview p { font-size: 16px; line-height: 1.6; }
.passport-overview .source-caption { font-size: 12px; }
.natural-fact, .natural-identification > article, .data-card, .catalog-branch { border: 0; border-radius: 12px; background: var(--site-panel); }
.passport-section { padding: 32px 0; border-top: 1px solid var(--site-line); }
.species-passport h2 { font-size: 24px; font-weight: 500; }
.species-passport .source-caption { color: var(--site-muted); }
.catalog-branches { padding: 24px 0 32px; }
.catalog-branch-grid { gap: 24px; }
.catalog-branch { overflow: hidden; padding: 0; }
.catalog-branch > a { display: flex; flex-direction: column; gap: 8px; padding: 16px 24px 8px; }
.catalog-branch > small { display: block; padding: 0 24px 20px; color: var(--site-muted); }
.catalog-branch img { width: 100%; height: 240px; object-fit: cover; }
.site-article-page .scientific-toc { grid-column: 2; grid-row: 1; align-self: start; background: var(--site-panel); border-radius: 12px; }
.site-design-header .reader-header-search { flex-direction: row-reverse; }
.site-design-header .reader-header-search input { order: 0; }
.site-design-header .reader-header-search input { display: block; }
@media (min-width: 1024px) {
  :root:not([data-navigation-expanded="false"]) .platform-sidebar .site-nav-group h2 { display: block; }
  :root:not([data-navigation-expanded="false"]) .platform-sidebar .site-nav-link {
    flex-direction: row; justify-content: flex-start; height: 28px; min-height: 28px; padding: 0 8px; margin: 0; gap: 8px;
  }
  :root:not([data-navigation-expanded="false"]) .platform-sidebar .site-nav-link strong { font-size: 14px; line-height: 16px; text-align: left; }
  .platform-sidebar .site-nav-icon img { width: auto; height: auto; max-width: 20px; max-height: 20px; }
  :root[data-navigation-expanded="false"] .platform-sidebar .site-nav-group h2 { display: none; }
  :root[data-navigation-expanded="false"] .platform-sidebar .site-nav-link { flex-direction: column; height: auto; min-height: 60px; gap: 4px; padding: 6px 0; text-align: center; }
  :root[data-navigation-expanded="false"] .platform-sidebar .site-nav-link strong { font-size: 10px; line-height: 1.25; text-align: center; }
}
@media (max-width: 1023px) {
  .site-home-heading h1 { font-size: 32px; }
  .site-home-groups { grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .site-home-actions { margin-top: 24px; }
  .site-home .primary, .site-home .secondary { min-width: 0; }
  .site-article-page .scientific-toc:not(.scientific-toc-desktop) { grid-column: 1; grid-row: auto; margin-bottom: 24px; }
  .site-article-page .scientific-body { grid-row: auto; }
}
@media (max-width: 650px) {
  .site-wordmark, .reader-header .brand .site-wordmark { width: 104px; height: auto; }
  .site-header-brand { flex-shrink: 0; }
  .site-home-heading { margin-bottom: 32px; }
  .site-home-grid, .site-home-feature, .site-home-research, .passport-intro { grid-template-columns: minmax(0, 1fr); }
  .site-home-groups { grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 12px; }
  .site-home-groups .site-home-card { padding: 16px; }
  .site-home-card { min-height: 172px; }
  .site-home-feature { gap: 24px; }
  .site-home-feature h3 { font-size: 28px; }
  .site-home .site-section { padding: 28px 0 32px; }
  .site-home-entry { margin-bottom: 32px; }
  .passport-title h1 { font-size: 30px; overflow-wrap: anywhere; }
  .passport-tabs { gap: 8px; flex-wrap: wrap; }
}
@media (max-width: 370px) { .site-wordmark, .reader-header .brand .site-wordmark { width: 88px; } }
:root[data-eg-theme="light"] .site-nav-icon img, :root[data-eg-theme="light"] .site-wordmark { filter: invert(1); }
@media (max-width: 1023px) { .site-nav-link, .reader-drawer .site-nav-link { min-height: 44px; height: auto; } }
@media (max-width: 370px) {
  .site-wordmark, .reader-header .brand .site-wordmark { width: 72px; }
  .site-languages.reader-languages { flex-basis: 64px; width: 64px; }
  .site-languages a { width: 28px; }
  .site-global-search, .site-design-header .reader-header-search { min-width: 32px; flex-basis: 32px; padding: 0 6px; }
  .site-header-account, .site-design-header .reader-header-login, .site-design-header [data-reader-account="header"] a { min-width: 48px; }
}
.site-catalog-table { padding: 12px 24px; border-radius: 12px; background: var(--site-panel); overflow-x: auto; }
.site-catalog-table table { border-collapse: collapse; width: 100%; min-width: 640px; }
.site-catalog-table thead th { padding: 16px 12px; color: var(--site-muted); font-size: 12px; font-weight: 400; text-align: left; }
.site-catalog-table tbody th, .site-catalog-table td { padding: 16px 12px; border-top: 1px solid var(--site-line); text-align: left; vertical-align: middle; font-size: 14px; font-weight: 400; }
.site-catalog-table tbody th { width: 32%; }
.site-catalog-table a { color: var(--site-text); text-decoration: none; }
.site-catalog-table a:hover { text-decoration: underline; }
.catalog-row-name { display: block; font-weight: 500; }
.site-catalog-table small { display: block; margin-top: 4px; color: var(--site-muted); font-size: 12px; }
.catalog-row-actions { display: flex; flex-direction: column; gap: 8px; font-size: 12px; }
.site-species-top { display: flex; align-items: center; justify-content: space-between; gap: 24px; flex-wrap: wrap; }
.site-species-top .passport-back { margin: 0; }
.site-species-actions { display: flex; align-items: center; gap: 8px; flex-wrap: wrap; }
.site-species-status { display: inline-flex; margin-top: 16px; padding: 12px 20px; border-radius: 8px; font-size: 12px; background: var(--site-accent); }
.site-species-overview { margin-bottom: 48px; }
.site-species-overview .passport-title h1 { font-style: normal; }
.site-species-overview .passport-title p { color: var(--site-muted); }
.site-species-overview .passport-tabs a { background: none; border: 0; border-radius: 0; padding: 12px 32px; text-decoration: none; color: var(--site-text); }
.site-species-overview .passport-tabs a:hover { box-shadow: inset 0 -2px var(--site-muted); }
.site-species-hero-photo { display: flex; align-items: center; justify-content: center; min-width: 0; border-radius: 12px; overflow: hidden; background: var(--site-panel); }
.site-species-hero-photo figure { margin: 0; width: 100%; border: 0; border-radius: 0; }
.site-species-hero-photo img { width: 100%; max-height: 390px; object-fit: contain; }
.site-species-hero-photo figcaption { padding: 12px 16px; font-size: 11px; line-height: 1.4; }
.site-species-overview .passport-overview { display: flex; flex-direction: column; align-items: flex-start; }
.site-species-overview .passport-overview .primary { margin-top: auto; }
.site-species-facts { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 24px; margin: 32px 0; }
.site-species-facts > article { background: var(--site-panel); border-radius: 12px; padding: 20px; }
.site-species-facts h3 { margin: 0 0 12px; font-size: 12px; font-weight: 400; color: var(--site-muted); }
.site-species-facts p { margin: 0; font-size: 16px; line-height: 1.5; }
.site-species-facts .source-caption { margin-top: 12px; font-size: 11px; color: var(--site-muted); }
.site-species-context { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); gap: 24px; }
.site-species-context > section { padding: 24px; border-radius: 12px; background: var(--site-panel); }
.site-species-context > section:last-child { background: var(--site-feature); }
.site-species-context h2 { font-size: 22px; font-weight: 500; margin: 0 0 20px; }
.site-species-context dl { display: grid; grid-template-columns: 1fr 2fr; gap: 12px; font-size: 14px; line-height: 1.5; }
.site-species-context dt { color: var(--site-muted); }
.site-species-context dd { margin: 0; }
.site-species-context p { color: var(--site-muted); font-size: 14px; }
.site-species-context a { color: var(--site-text); font-size: 14px; }
.site-species-context > section:last-child a { display: block; padding: 12px 16px; margin-top: 8px; border-radius: 8px; background: var(--site-panel); text-decoration: none; }
.site-species-page #species-description { padding-top: 32px; border-top: 1px solid var(--site-line); scroll-margin-top: 100px; }
@media (max-width: 650px) {
  .site-catalog-table { padding: 8px; }
  .site-species-facts, .site-species-context { grid-template-columns: minmax(0, 1fr); gap: 12px; }
  .site-species-top { align-items: flex-start; gap: 12px; }
  .site-species-top .passport-back, .site-species-actions { width: 100%; }
  .site-species-actions { gap: 8px; }
  .site-species-actions > * { flex: 1; }
  .site-species-actions a { display: flex; justify-content: center; align-items: center; min-height: 44px; padding: 8px; font-size: 12px; text-align: center; }
  .site-species-overview .passport-tabs { display: grid; grid-template-columns: repeat(2, minmax(0, 1fr)); }
  .site-species-overview .passport-tabs a { padding: 12px 8px; text-align: center; }
}

.platform-sidebar:has(.site-nav-link) .platform-sidebar-footer { display: flex; align-items: center; justify-content: space-between; gap: 8px; min-height: 40px; padding: 4px 12px; margin-top: 4px; }

/* The React theme uses 100/200/400 for header, drawer and modal. */
.site-public-app .site-design-header { z-index: 100; }

.reader-header .reader-drawer > nav { flex-wrap: nowrap; align-items: stretch; }
.reader-drawer > nav > * { flex-shrink: 0; }
.reader-header .reader-drawer nav .site-nav-link { min-height: 44px; height: auto; padding: 8px; }

/* Preserve contrast against the same dark controls in both colour themes. */
.site-design-header .site-icon-button:not(.site-header-help), .site-design-header .reader-desktop-menu, .site-design-header .reader-menu > summary { color: var(--eco-primitive-neutral-white); }
.site-design-header .site-languages.reader-languages a { border: 0; min-height: 32px; padding: 0; color: var(--eco-primitive-neutral-light-gray); }
.site-design-header .site-languages.reader-languages a[aria-current="true"] { color: var(--eco-primitive-neutral-white); }
.platform-sidebar .site-nav-link strong, .reader-drawer .site-nav-link strong { color: inherit; }
.site-species-status { color: var(--eco-primitive-neutral-white); }
:root[data-eg-theme="light"] .site-nav-link[aria-current="page"] .site-nav-icon img { filter: none; }
.platform-sidebar:has(.site-nav-link) .platform-sidebar-footer .reader-hide-menu { margin: 0; }